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A hidden diversity of ceratopsian dinosaurs in Late Cretaceous Europe

A hidden diversity of ceratopsian dinosaurs in Late Cretaceous Europe

  • Csiki-Sava, Z., Buffetaut, E., Ősi, A., Pereda-Suberbiola, X. & Brusatte, S. L. Island life in the Cretaceous — faunal composition, biogeography, evolution, and extinction of land-living vertebrates on the Late Cretaceous European archipelago. Zookeys 469, 1–161 (2015).

    Article 

    Google Scholar 

  • Weishampel, D. B., Jianu, C.-M., Csiki, Z. & Norman, D. B. Osteology and phylogeny of Zalmoxes (n. g.), an unusual euornithopod dinosaur from the latest Cretaceous of Romania. J. Syst. Palaeontol. 1, 65–123 (2003).

    Article 

    Google Scholar 

  • Augustin, F., Ősi, A. & Csiki-Sava, Z. The Rhabdodontidae (Dinosauria, Ornithischia), an enigmatic dinosaur group endemic to the Late Cretaceous European Archipelago. Fossil Rec. 26, 171–189 (2023).

    Article 

    Google Scholar 

  • Czepiński, Ł & Madzia, D. Osteology, phylogenetic affinities, and palaeobiogeographic significance of the bizarre ornithischian dinosaur Ajkaceratops kozmai from the Late Cretaceous European archipelago. Zoo. J. Linn. Soc. 202, 1–25 (2024).

    Article 

    Google Scholar 

  • Ősi, A., Butler, R. J. & Weishampel, D. B. A Late Cretaceous ceratopsian dinosaur from Europe with Asian affinities. Nature 465, 466–468 (2010).

    Article 
    ADS 
    PubMed 

    Google Scholar 

  • Ryan, M. J., Chinnery-Allgeier, B. J. & Eberth, D. A. New Perspectives on Horned Dinosaurs (Indiana Univ. Press, 2010).

  • You, H. & Dodson, P. in The Dinosauria (eds Weishampel, D. B., Dodson, P. and Osmólska, H.) 478–493 (Univ. California Press, 2004).

  • Norman, D. B., Sues, H.-D., Witmer, L. M. & Coria, R. A. in The Dinosauria (eds Weishampel, D. B., Dodson, P. and Osmólska, H.) 393–412 (Univ. California Press, 2004).

  • Maidment, S. C. R. & Barrett, P. M. Does morphological convergence imply functional similarity? A test using the evolution of quadrupedalism in ornithischian dinosaurs. Proc. R. Soc. B 279, 3765–3771 (2012).

    Article 
    PubMed 
    PubMed Central 

    Google Scholar 

  • Nabavizadeh, A. Evolutionary trends in the jaw adductor mechanics of ornithischian dinosaurs. Anat. Rec. 299, 271–294 (2016).

    Article 

    Google Scholar 

  • Virág, A. & Ősi, A. Morphometry, microstructure, and wear pattern of neornithischian dinosaur teeth from the Upper Cretaceous Iharkút locality (Hungary). Anat. Rec. 300, 1439–1463 (2017).

    Article 

    Google Scholar 

  • Farke, A. A., Maxwell, W. D., Cifelli, R. L. & Wedel, M. J. A ceratopsian dinosaur from the Lower Cretaceous of western North America, and the biogeography of Neoceratopsia. PLoS ONE 9, e112055 (2014).

    Article 
    ADS 
    PubMed 
    PubMed Central 

    Google Scholar 

  • Longrich, N. R. A ceratopsian dinosaur from the Late Cretaceous of eastern North America, and implications for dinosaur biogeography. Cret. Res. 57, 199–207 (2016).

    Article 

    Google Scholar 

  • Morschhauser, E. M. et al. Phylogenetic history of Auroraceratops rugosus (Ceratopsia: Ornithischia) from the Lower Cretaceous of Gansu Province, China. J. Vertebr. Paleontol. 38, 117–147 (2019).

    Article 

    Google Scholar 

  • Godefroit, P. & Lambert, O. A re-appraisal of Craspedodon lonzeensis Dollo, 1883 from the Upper Cretaceous of Belgium: the first record of a neoceratopsian dinosaur in Europe? Bull. l’Inst. R. Sci. Naturelles Belgique Sci. Terre 77, 83–93 (2007).

    Google Scholar 

  • Lindgren, J. et al. The first neoceratopsian dinosaur remains from Europe. Palaeontology 50, 929–937 (2007).

    Article 

    Google Scholar 

  • Han, F., Forster, C. A., Xu, X. & Clark, J. M. Postcranial anatomy of Yinlong downsi (Dinosauria: Ceratopsia) from the Upper Jurassic Shishugou Formation of China and the phylogeny of basal ornithischians. J. Syst. Palaeontol. 16, 1159–1187 (2018).

    Article 

    Google Scholar 

  • Butler, R. J. & Sullivan, R. M. The phylogenetic position of Stenopelix valdensis from the Lower Cretaceous of Germany and the early fossil record of Pachycephalosauria. Acta Palaeontol. Pol. 54, 21–34 (2009).

    Article 

    Google Scholar 

  • Bertozzo, F., Kecheng, N., Gillette, N. V. & Godefroit, P. Anatomical description and digital reconstruction of the skull of Jeholosaurus shangyuanensis (Dinosauria, Ornithopoda) from China. PLoS ONE 20, e0312519 (2025).

    Article 
    PubMed 
    PubMed Central 
    CAS 

    Google Scholar 

  • Matheron, P. in Mémoires de l’Académie Impériale des Sciences, Belles – Lettres et Arts de Marseille 1–39 (Académie des Sciences, Lettres et Arts de Marseille, Marseille, 1869).

  • Pincemaille-Quillevere, M. Description d’un squelette partiel de Rhabdodon priscus (Euornithopoda) du Crétacé supérieur de Vitrolles (Bouches du Rhône, France). Oryctos 4, 39–70 (2002).

    Google Scholar 

  • Ősi, A., Prondvai, E., Butler, R. J. & Weishampel, D. B. Phylogeny, histology and inferred body size evolution in a new rhabdodontid dinosaur from the Late Cretaceous of Hungary. PLoS ONE 7, e44318 (2012).

    Article 
    ADS 
    PubMed 
    PubMed Central 

    Google Scholar 

  • Magyar, J., Csiki-Sava, Z., Ősi, A., Augustin, F. J. & Botfalvai, G. Rhabdodontid (Dinosauria, Ornithopoda) diversity suggested by the first documented occurrence of associated cranial and postcranial material at Vălioara (uppermost Cretaceous Densuș-Ciula Formation, Hațeg Basin, Romania). Cret. Res. 156, 105810 (2024).

    Article 

    Google Scholar 

  • Czepiński, Ł. Ontogeny and variation of a protoceratopsid dinosaur Bagaceratops rozhdestvenskyi from the Late Cretaceous of the Gobi Desert. Hist. Biol. 32, 1394–1421 (2020).

    Article 

    Google Scholar 

  • Gilmore, C. W. Osteology of the Jurassic reptile Camptosaurus with a revision of the species of the genus and descriptions of two new species. Proc. US Nat. Mus. 36, 197–302 (1909).

    Article 

    Google Scholar 

  • Thomas, D. A. The cranial anatomy of Tenontosaurus tilletti Ostrom, 1970 (Dinosauria, Ornithopoda). Palaeontol. Electron. 18.2.37A, 1–99 (2015).

  • Porro, L. B., Witmer, L. M. & Barrett, P. M. Digital preparation and osteology of the skull of Lesothosaurus diagnosticus (Ornithischia: Dinosauria). PeerJ 3, e1494 (2015).

    Article 
    PubMed 
    PubMed Central 

    Google Scholar 

  • Horner, J. R. & Goodwin, M. B. Ontogeny of cranial epi-ossifications in Triceratops. J. Vertebr. Paleontol. 28, 134–144 (2008).

    Article 

    Google Scholar 

  • Loewen, M. A. et al. Lokiceratops rangiformis gen. et sp. nov. (Ceratopsidae: Centrosaurinae) from the Campanian Judith River Formation of Montana reveals rapid regional radiations and extreme endemism within centrosaurine dinosaurs. PeerJ 12, e17224 (2024).

    Article 
    PubMed 
    PubMed Central 

    Google Scholar 

  • Longrich, N. R. & Field, D. J. Torosaurus is not Triceratops: ontogeny in chasmosaurine ceratopsids as a case study in dinosaur taxonomy. PLoS ONE 7, e32623 (2012).

    Article 
    ADS 
    PubMed 
    PubMed Central 
    CAS 

    Google Scholar 

  • Tykoski, R. S., Fiorillo, A. R. & Chiba, K. New data and diagnosis for the Arctic ceratopsid dinosaur Pachyrhinosaurus perotorum. J. Syst. Palaeontol. 17, 1397–1416 (2019).

    Article 

    Google Scholar 

  • Zhao, X., Cheng, Z. & Xu, X. The earliest ceratopsian from the Tuchengzi Formation of Liaoning, China. J. Vertebr. Paleontol. 19, 681–691 (2009).

    Google Scholar 

  • Sereno, P. C. in New Perspectives on Horned Dinosaurs (Ryan, M. J., Chinnery-Allgeier, B. J. & Eberth, D. A.) 21–58 (Indiana Univ. Press, 2010).

  • Maryańska, T. & Osmólska, H. Protoceratopsidae (Dinosauria) of Asia. Palaeontol. Pol. 33, 133–181 (1975).

    Google Scholar 

  • You, H. & Dong, Z. A new protoceratopsid (Dinosauria: Neoceratopsia) from the Late Cretaceous of Inner Mongolia, China. Acta Geol. Sin. 77, 299–303 (2003).

    Article 

    Google Scholar 

  • Wolfe, D. G. et al. in New Perspectives on Horned Dinosaurs (Ryan, M. J., Chinnery-Allgeier, B. J. & Eberth, D. A.) 91–98 (Indiana Univ. Press, 2010).

  • Kirkland, J. I. & DeBlieux, D. D. in New Perspectives on Horned Dinosaurs (Ryan, M. J., Chinnery-Allgeier, B. J. & Eberth, D. A.) 117–140 (Indiana Univ. Press, 2010).

  • Han, F., Forster, C. A., Clark, J. M. & Xu, X. Cranial anatomy of Yinlong downsi (Ornithischia: Ceratopsia) from the Upper Jurassic Shishugou Formation of Xinjiang, China. J. Vertebr. Paleontol. 36, e1029579 (2016).

    Article 

    Google Scholar 

  • Morschhauser, E. M., Li, D., You, H. & Dodson, P. Cranial anatomy of the basal neoceratopsian Auroraceratops rugosus (Ornithischia: Ceratopsia) from the Yujingzi Basin, Gansu Province, China. J. Vertebr. Paleontol. 38, 36–68 (2019).

    Article 

    Google Scholar 

  • Hübner, T. & Rauhut, O. W. M. A juvenile skull of Dysalotosaurus lettowvorbecki (Ornithischia: Iguanodontia), and implications for cranial ontogeny, phylogeny, and taxonomy in ornithopod dinosaurs. Zoo. J. Linn. Soc. 160, 366–396 (2010).

    Article 

    Google Scholar 

  • Galton, P. M. The ornithischian dinosaur Hypsilophodon from the Wealden of the Isle of Wight. Bull. Brit. Mus. Nat. Hist. Geol. 25, 1–152 (1974).

    Google Scholar 

  • Brown, B. Leptoceratops, a new genus of Ceratopsia from the Edmonton Cretaceous of Alberta. Bull. Am. Mus. Nat. Hist. 33, 567–580 (1914).

    Google Scholar 

  • Makovicky, P. J. & Norell, M. A. Yamaceratops dorngobiensis, a new primitive ceratopsian (Dinosauria: Ornithischia) from the Cretaceous of Mongolia. Am. Mus. Novit. 3530, 1–42 (2006).

    Article 

    Google Scholar 

  • Tanoue, K., Grandstaff, B. S., You, H. L. & Dodson, P. Jaw mechanics in basal Ceratopsia. Anat. Rec. 292, 1352–1369 (2009).

    Article 

    Google Scholar 

  • Butler, R. J., Upchurch, P. & Norman, D. B. The phylogeny of the ornithischian dinosaurs. J. Syst. Palaeontol. 6, 1–40 (2008).

    Article 

    Google Scholar 

  • Dodson, P., Forster, C. A. & Sampson, S. A. in The Dinosauria (eds Weishampel, D. B., Dodson, P. & Osmólska. H.) 494–516 (Univ. California Press, 2004).

  • Dieudonné, P.-E., Cruzado-Caballero, P., Godefroit, P. & Tortosa, T. A new phylogeny of cerapodan dinosaurs. Hist. Biol. 33, 2335–2355 (2021).

    Article 

    Google Scholar 

  • Maidment, S. C. R. & Barrett, P. M. Osteological correlates for quadrupedality in ornithischian dinosaurs. Acta Palaeontol. Pol. 59, 53–70 (2014).

    Google Scholar 

  • Chanthasit, P. The ornithopod dinosaur Rhabdodon from the Late Cretaceous of France: anatomy, systematics and paleobiology. PhD Thesis: Université Claude Bernard (2010).

  • Csiki, Z., Vremir, M., Brusatte, S. L. & Norell, M. A. An aberrant island-dwelling theropod dinosaur from the Late Cretaceous of Romania. Proc. Natl Acad. Sci. USA 107, 15357–15361 (2010).

    Article 
    ADS 
    PubMed 
    PubMed Central 
    CAS 

    Google Scholar 

  • Wolfe, D. G. & Kirkland, J. I. Zuniceratops christopheri n. gen & n. sp., a ceratopsian dinosaur from the Moreno Hill Formation (Cretaceous, Turonian) of west-central New Mexico. New Mex. Mus. Nat. Hist. Sci. Bull. 14, 303–317 (1998).

    Google Scholar 

  • Sues, H. D. & Averianov, A. Turanoceratops tardabilis — the first ceratopsid dinosaur from Asia. Naturwissenschaften 96, 645–652 (2009).

    Article 
    ADS 
    PubMed 
    CAS 

    Google Scholar 

  • Herbst, E. C., Meade, L. E., Lautenschlager, S., Fioritti, N. & Scheyer, T. M. A toolbox for the retrodeformation and muscle reconstruction of fossil specimens in Blender. Roy. Soc. Open Sci. 9, 220519 (2022).

    Article 
    ADS 

    Google Scholar 

  • Goloboff, P. A. & Catalano, S. A. TNT version 1.5, including a full implementation of phylogenetic morphometrics. Cladistics 32, 221–238 (2016).

    Article 
    PubMed 

    Google Scholar 

  • Maddison, W. P. & Maddison, D. R. Mesquite: a modular system for evolutionary analysis. Version 3.04. Mesquite https//www.mesquiteproject.org (2025).

  • Ronquist, F. et al. MrBayes 3.2: efficient Bayesian phylogenetic inference and model choice across a large model space. Syst. Biol. 61, 539–542 (2012).

    Article 
    PubMed 
    PubMed Central 

    Google Scholar 

  • Lewis, P. O. A likelihood approach to estimating phylogeny from discrete morphological character data. Syst. Biol. 50, 913–925 (2001).

    Article 
    PubMed 
    CAS 

    Google Scholar 

  • Nylander, J. A. A., Ronquist, F., Huelsenbeck, J. & Nieves-Aldrey, J. L. Bayesian phylogenetic analysis of combined data. Syst. Biol. 53, 47–67 (2004).

    Article 
    PubMed 

    Google Scholar 

  • Kass, R. E. & Raftery, A. E. Bayes factor. J. Am. Stat. Assoc. 90, 773–795 (1995).

    Article 
    MathSciNet 
    MATH 

    Google Scholar 

  • Maidment, S. C. R., Butler, R. J., Brusatte, S. L., Meade, L. E., Augustin, F. J.,Csiki-Sava, Z. & Ősi, A. Data for “A hidden diversity of ceratopsian dinosaurs in Late Cretaceous Europe”. Zenodo https://doi.org/10.5281/zenodo.17098636 (2025)

  • Słowiak, J., Tereshchenko, V. S. & Fostowicz-Frelik, Ł Appendicular skeleton of Protoceratops andrewsi (Dinosauria: Ornithischia): comparative morphology, ontogenetic changes, and implications for non-ceratopsid ceratopsian locomotion. PeerJ 7, e7324 (2019).

    Article 
    PubMed 
    PubMed Central 

    Google Scholar 


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